The impact of grazing on spatial variability of humus profile properties in a grass-encroached Scots pine ecosystem

Grazing is often used in management of grass-encroached Scots pine (Pinus sylvestris) forests in The Netherlands, because it is thought to increase diversity of vegetation and humus profiles. To test this hypothesis, two 1 ha plots in a primary pine stand on drift sand were grazed for 7 years at high and moderate intensities. The humus profiles of these two sites and of a control site were sampled at various scales to assess changes in grass biomass, thickness of ectorganic horizons, organic matter stocks, root stocks, and pH. Whether grazed or not, site factors such as slope angle and aspect, distance to trees and parent material did not explain the spatial variability of the plots. Spatial variability could be linked to the tussock structure of the grass layer. With continued grazing, ectorganic horizons became more compact (particularly the F horizon), stocks of organic matter increased, grass biomass decreased, variances tended to decrease and ranges tended to increase. These results suggest that at the landscape scale, where much variation in grazing intensity can be expected, spatial variability of humus profile properties and thus site diversity will increase. However, grazing intensities in the experiment were far greater than those normally employed in management of grass-encroached forests. Therefore grazing will scarcely affect site diversity, unless grazing variability is enhanced by special treatments, such as temporary fencing to create locally high grazing intensities.