Deep-sea trace fossils from the mixed carbonate-siliciclastic flysch of the Monte Antola Formation (Late Campanian-Maastrichtian), North Apennines, Italy

Twenty one ichnogenera and 30 ichnospecies are recognized in the Late Campanian – Maastrichtian Monte Antola Formation of Italy, most of them for the first time. They belong to a specific mixture of the Nereites and Ophiomorpha rudis ichnosubfacies of the Nereites ichnofacies. Composition and abundance of trace fossils vary from outcrop to outcrop. Chondrites intricatus, Ch. targionii, Nereites irregularis, Cladichnus fischeri and Trichichnus linearis are most common. Chondrites patulus, Ophiomorpha recta and “Spirophycus” bicornis trace fossils are less common in outcrops, and the other trace fossils are rare or very rare. Generally, ichnogeneric diversity in Cretaceous–Cainozoic calcareous and mixed siliciclastic-calcareous flysch deposits is distinctly lower than in siliciclastic flysch. Significantly, the foremost pre-depositional forms are distinctly rare, probably due to preservational and depositional factors.