Potential freshwater dinocysts from marine upper Cenomanian to upper Coniacian strata of Tarfaya, northwest Africa: Three new species of Bosedinia

Three new species of Bosedinia He emend. Chen et al. are described: B. alveolata, B. spinosa and B. tarfayensis. The diagnosis of the genus Bosedinia is emended to include forms with an alveolate wall and/or hair-like spiny processes. The appearance in massive blooms across considerable section intervals closely related to and within oceanic anoxic event 2 (Cenomanian–Turonian) and anoxic event 3 (Coniacian–Santonian) suggests episodic salinity stratification as important aspect for enhanced organic carbon accumulation.