Fire-related landform associations of remnant old-growth trees in the southern Washington Cascade Range

The spatial distribution of biological legacies left by natural disturbances is an important source of variability in forest development. We investigated one type of biological legacy: remnant old-growth trees persisting in mature Douglas-fir forests. We hypothesized that persistence varies with topographic heterogeneity influencing fire behavior. Our two study areas are located in the southern Washington Cascade Range, USA. They have an unfragmented, mature forest cover that regenerated following wildfire. We mapped all remnant old-growth trees (live and dead) within 4.2-6.4 km long belt transects. Digital elevation models were used to generate convergent and divergent landform classes. Frequency analysis was used to test for landform associations. Live remnant western hemlock and western redcedar were strongly associated with convergent landforms and aspects that had greater availability of soil moisture. Live remnant Douglas-fir were most abundant, but were not correlated with convergence or divergence, although certain landforms had higher concentrations. Remnant snags were abundant across convergent and divergent landforms. We conclude that species with low fire resistance survive most frequently on landforms that have a dampening effect on fire intensity. Topographic variability may indirectly influence ecological functions associated with biological legacies by affecting the spatial distributions of remnant old-growth trees.